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Description
The ability of Neandertals to cope with the oscillating climate of the late Pleistocene and the extent to which these climate changes affected local Neandertal habitats remain unanswered anthropological topics of considerable scientific interest. Understanding the impact of climatic instability on Neandertals is critical for reconstructing the behaviors of our

The ability of Neandertals to cope with the oscillating climate of the late Pleistocene and the extent to which these climate changes affected local Neandertal habitats remain unanswered anthropological topics of considerable scientific interest. Understanding the impact of climatic instability on Neandertals is critical for reconstructing the behaviors of our closest fossil relatives and possibly identifying factors that contributed to their extinction. My work aimed to test the hypotheses that 1) cold climates stressed Neandertal populations, and 2) that global climate changes affected local Neandertal habitats. An analysis of Neandertal butchering on Cervus elaphus, Rangifer tarandus, and Capreolus capreolus skeletal material deposited during global warm and cold phases from two French sites - Pech de l'Azé IV and Roc de Marsal - was conducted to assess the impact of climate change on butchering strategies and resource extraction. Results from a statistical analysis of surface modification on all marrow yielding long bones, including the 1st phalanx, demonstrated that specimens excavated from the cold levels at each cave have more cut marks (Wald χ2= 51.33, p= <0.001) and percussion marks (Wald χ2= 4.92, p= 0.02) than specimens from the warm levels after controlling for fragment size. These results support the hypothesis that Neandertals were nutritionally stressed during glacial cycles. The hypothesis that global climates affected local habitats was tested through radiogenic strontium isotopic reconstruction of large herbivore mobility patterns (e.g., Bison, Equus, Cervus and Rangifer), because it is known that in the northern hemisphere, mammals migrate less in warm, well-vegetated environments, but more in cold, open environments. Identifying isotopic variation in mammalian fossils enables mobility patterns to be inferred, providing an indication of whether environments at Pech de l'Azé IV and Roc de Marsal tracked global climates. Results from this study indicate that Neandertal prey species within the Dordogne Valley of France did not undertake long distance round-trip migrations in glacial or interglacial cycles, maintaining the possibility that local habitats did not change in differing climatic cycles. However, because Neandertals were nutritionally stressed the most likely conclusion is that glacial cycles decreased herbivore populations, thus stressing Neandertals.
ContributorsHodgkins, Jamie Melichar (Author) / Marean, Curtis W (Thesis advisor) / Reed, Kaye E (Thesis advisor) / Knudson, Kelly J. (Committee member) / Spencer, Lillian M (Committee member) / Arizona State University (Publisher)
Created2012
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Description
In recent years, southern Africa has figured prominently in the modern human origins debate due to increasing evidence for precocious behaviors considered to be unique to our species. These significant findings have included bone tools, shell beads, engraved ostrich eggshell, and heavily ground and engraved ochre fragments. The presence of

In recent years, southern Africa has figured prominently in the modern human origins debate due to increasing evidence for precocious behaviors considered to be unique to our species. These significant findings have included bone tools, shell beads, engraved ostrich eggshell, and heavily ground and engraved ochre fragments. The presence of ochre in Middle Stone Age (MSA, ~250-40kya) archaeological sites in southern Africa is often proposed as indirect evidence for the emergence of symbolic or artistic behavior, a uniquely modern human trait. However, there is no remaining artwork from this period and there is significant debate about what the ochre may have been used for. With a few exceptions, ochre has gone largely unstudied. This project tested competing models for ochre use within the Pinnacle Point (PP), South Africa research area. Combined results from characterization and sourcing analyses, color classification, heat treatment analysis, and hafting experiments suggest MSA ochre is tied to early symbolic or ritual behavior.
ContributorsBernatchez, Jocelyn A (Author) / Marean, Curtis W (Thesis advisor) / Bearat, Hamdallah (Committee member) / Abbott, David (Committee member) / Arizona State University (Publisher)
Created2012
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Description
Two of the defining behaviors associated with the hominin lineage are an increased reliance on tool use and the routine incorporation of animal tissue in the diet. These adaptations have been linked to numerous downstream consequences including key physiological adaptations as well as social and cognitive effects associated with modern

Two of the defining behaviors associated with the hominin lineage are an increased reliance on tool use and the routine incorporation of animal tissue in the diet. These adaptations have been linked to numerous downstream consequences including key physiological adaptations as well as social and cognitive effects associated with modern humans. Thus, a critical issue in human evolution is how to determine when hominins began incorporating significant amounts of meat into their diets. Bone surface modifications (BSM) have long been recognized as a powerful inferential tool in identifying the differential involvement of actors responsible for altering assemblages of bone recovered from both archaeological and paleontological contexts and remain a primary source of direct evidence for butchery activities. Thus, determining the spatiotemporal context of increased carnivory in the hominin lineage relies on the accurate identification of fossil BSM.

Multidecade-long debates over the agents responsible for individual BSM indicate systemic flaws in historical approaches to identification. These debates are in part due to the extreme morphological overlap between BSM produced by certain agents of modification. The primary goal of this dissertation project therefore, is to construct probability models of BSM capable of identifying individual marks with an associated probability of assignment. Using a multivariate Bayesian approach to analyze experimentally-generated BSM data, this dissertation uses two different models, one incorporating both two and three-dimensional (3D) metric and attribute data associated with individual BSM and a second model comparing 3D geometric morphometric (GM) shape data associated with BSM.

The 2D/3D attribute model of BSM is used evaluate an assemblage of fossil BSM recovered from the Ledi-Geraru research area, Ethiopia (2.82 Ma) in spatiotemporal association with early Homo. The results of the analysis reveal compelling evidence for early butchery activities, suggesting hominins may have been using both modified and unmodified stone implements to process carcasses.

The second model, based upon 3D GM data, was used to evaluate the earliest purported evidence for stone-mediated butchery at Dikika, Ethiopia (3.39 Ma). The Dikika marks have been argued to be the result of crocodile feeding, trampling, and butchery by three different research groups. The 3D GM model evaluates the likelihood of each of these actors in the production of the controversial Dikika marks.
ContributorsHarris, Jacob A (Author) / Marean, Curtis W (Thesis advisor) / Hill, Kim (Committee member) / Boyd, Robert (Committee member) / Thompson, Jessica (Committee member) / Campisano, Christopher (Committee member) / Arizona State University (Publisher)
Created2019
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Description
This dissertation research describes the hunting behavior of early modern humans through the analysis of vertebrate faunal remains from Contrebandiers Cave, Morocco. Contrebandiers Cave is located in the town of Témara and is roughly 250 meters from the current shoreline of the Atlantic Ocean. The cave was excavated in the

This dissertation research describes the hunting behavior of early modern humans through the analysis of vertebrate faunal remains from Contrebandiers Cave, Morocco. Contrebandiers Cave is located in the town of Témara and is roughly 250 meters from the current shoreline of the Atlantic Ocean. The cave was excavated in the 1950s and 1970s by l’Abbé Roche, and again starting in 2007 by Dibble and El Hajraoui with total station plotting of finds. Contrebandiers Cave contains Middle Stone Age (MSA) deposits dated to Marine Isotope Stages (MIS) 5e, 5d and 5c, ~120,000 to ~96,000 years ago. The Later Stone Age (LSA) deposits are dated to MIS 2, ~20,000 years ago. The entirety of the ~12,000 vertebrate faunal remains from Dibble and El Hajraoui’s excavation were analyzed for taxonomic and taphonomic identification.

A total of 67 vertebrate taxa were identified and include ungulates, carnivores, lagomorphs, birds, tortoises, snakes and fish. The faunal remains from Contrebandiers Cave preserve surface modification that indicates both humans and carnivores acted as agents of prey accumulation. Skeletal element representation and surface modification of ungulate remains suggest that humans had primary access to small, medium and large-bodied prey. In the MSA levels, carnivore skeletal remains preserve surface modification that is interpreted as being indicative of behavior associated with skinning for fur removal.

The vertebrate faunal remains from MIS 5e and 5d indicate that humans were hunting grazers and mixed feeders from open habitats and suids from mixed habitats. The faunal remains from MIS 5c indicate that humans focused less on suids and more on mixed feeders from open habitats. The vertebrate faunal remains from MIS 2 reveal humans hunting grazers from dry, open habitats. This research provides a description of human hunting behavior in North Africa, and contributes to our understanding of early modern human behavior prior to dispersal out of Africa.
ContributorsHallett, Emily Yuko (Author) / Marean, Curtis W (Thesis advisor) / Reed, Kaye E (Committee member) / Dibble, Harold L. (Committee member) / Arizona State University (Publisher)
Created2018
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Description
Climate and environmental forcing are widely accepted to be important drivers of evolutionary and ecological change in mammal communities over geologic time scales. This paradigm has been particularly influential in studies of the eastern African late Cenozoic fossil record, in which aridification, increasing seasonality, and C4 grassland expansion are seen

Climate and environmental forcing are widely accepted to be important drivers of evolutionary and ecological change in mammal communities over geologic time scales. This paradigm has been particularly influential in studies of the eastern African late Cenozoic fossil record, in which aridification, increasing seasonality, and C4 grassland expansion are seen as having shaped the major patterns of human and faunal evolution. Despite the ubiquity of studies linking climate and environmental forcing to evolutionary and ecological shifts in the mammalian fossil record, many central components of this paradigm remain untested or poorly developed. To fill this gap, this dissertation employs biogeographical and macroecological analyses of present-day African mammal communities as a lens for understanding how abiotic change may have shaped community turnover and structure in the eastern African Plio-Pleistocene. Three dissertation papers address: 1) the role of ecological niche breadth in shaping divergent patterns of macroevolutionary turnover across clades; 2) the effect of climatic and environmental gradients on community assembly; 3) the relative influence of paleo- versus present-day climates in structuring contemporary patterns of community diversity. Results of these papers call into question many tenets of current theory, particularly: 1) that niche breadth differences (and, by extension, their influence on allopatric speciation) are important drivers of macroevolution, 2) that climate is more important than biotic interactions in community assembly, and 3) that communities today are in equilibrium with present-day climates. These findings highlight the need to critically reevaluate the role and scale-dependence of climate in mammal evolution and community ecology and to carefully consider potential time lags and disequilibrium dynamics in the fossil record.
ContributorsRowan, John (Author) / Reed, Kaye E (Thesis advisor) / Campisano, Christopher J (Committee member) / Franklin, Janet (Committee member) / Marean, Curtis W (Committee member) / Arizona State University (Publisher)
Created2018