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Description
Identifying the ecological role, or niche, that a species occupies within their larger community elucidates environmental adaptability and evolutionary success. This dissertation investigates the occupied niche of chimpanzees (Pan troglodytes schweinfurthii) living in an open, dry savanna-woodland environment by examining patterns of resource use and interspecific interactions. Data were collected

Identifying the ecological role, or niche, that a species occupies within their larger community elucidates environmental adaptability and evolutionary success. This dissertation investigates the occupied niche of chimpanzees (Pan troglodytes schweinfurthii) living in an open, dry savanna-woodland environment by examining patterns of resource use and interspecific interactions. Data were collected October 2010--November 2011 at Issa, in the Ugalla region of western Tanzania, which is one of the driest, most open, and seasonal habitats inhabited by chimpanzees. Unlike most primatological studies which employ methods that include focal follows, this study focused instead on observing 'resource patches' for chimpanzees. Patch focals allow for the observation of all animals within a study area; capture resources that are not used by the study species; and are particularly well suited for unhabituated communities. In order to better understand relationships between environment and behavior, data collected at Issa are compared with published data from other chimpanzee populations. Issa chimpanzees were expected to have broader resource use than forest chimpanzees, as well as increased competition with other fauna, due to fewer available resources. However, in contrast to the assumption of food scarcity in dry habitats, dietary resources were available throughout the year. Like other populations, the diet of Issa chimpanzees consisted of mostly fruit, but unlike at other sites, the majority of plants consumed were woodland species. Additionally, although chimpanzees and other fauna shared spatial and dietary resources, there was only nominal overlap. These results point to extremely low levels of indirect competition between chimpanzees and other fauna. Despite extensive study of forest chimpanzees, little is known about their role within their faunal community in open, dry habitats, nor about how greater seasonality affects resource use. This project addresses both of these important issues and fosters novel approaches in anthropological studies, especially in reference to chimpanzee ecology and evolution. Understanding current chimpanzee behavioral relationships with their environments shapes hypotheses about their pasts, and also informs predictions about behaviors of similar taxa in paleo-environments. Lastly, examining the ecological role of chimpanzees within their larger communities will influence the formation of, as well as evaluate, conservation strategies.
ContributorsRussak, Samantha M (Author) / Reed, Kaye E (Thesis advisor) / Nash, Leanne T. (Committee member) / Schwartz, Gary T (Committee member) / Arizona State University (Publisher)
Created2013
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Description
Unanswered questions about the evolution of human gender abound and are salient across the anthropological disciplines and beyond. Did adult sex-typed behavioral tendencies actually evolve? If so, when? For what purpose? The best way to gain insight into the evolution of human gender is to understand the evolution and development

Unanswered questions about the evolution of human gender abound and are salient across the anthropological disciplines and beyond. Did adult sex-typed behavioral tendencies actually evolve? If so, when? For what purpose? The best way to gain insight into the evolution of human gender is to understand the evolution and development of sex-typed behavior in comparative primate taxa. Captive research indicates that there are many proximate factors likely to shape the development of sex-typed behavior in non-human primates—prenatal and postnatal endocrinological experience, social experience, ecological factors, and their interactions. However, it is largely unknown how sex-typed behavior proceeds and is shaped by those factors in evolutionarily salient environments. This study investigated one—whether extrinsic sexually differentiated social interactions are likely influential in the development of adult sex-typed behavior in wild-living Lemur catta. Little is known about sex-typed development in this species or in strepsirrhines in general. This research therefore addresses an important phylogenetic gap in our understanding of primate sex-typed development. Behavioral observations were carried out on mixed cross-sectional sample of adult females (n=10), adult males (n=8), yearling females (n=4), yearling males (n=4), and newborn females (n=16) and males (n=14) at Beza Mahafaly Special Reserve in southwest Madagascar from September 2008 to August 2009. Twenty-three sex-typed behaviors were identified in adults using linear mixed effects models and models of group response profiles through time. Of those, only eight had a pre-pubertal developmental component. Infants did not exhibit any sex differences in behavior, but juveniles (prepubertal, weaned individuals) resembled adults in their (relatively few) patterns of expression of sex-typed behavior. Most adult sex-typed behaviors in this species apparently develop at or after puberty and may be under gonadal hormone control. Those that develop before puberty do not likely depend on extrinsic sexually differentiation social interactions for their development, because there is no clear evidence that infants and juvenile male and females are not treated differently by others according to sex. If sexually differentiated social interactions are important for sex-typed behavioral development in subadult ,italic>Lemur catta, they are likely intrinsically (rather than extrinsically) driven.
ContributorsMeredith, Stephanie Lynn (Author) / Nash, Leanne T. (Thesis advisor) / Reed, Kaye E (Committee member) / Schwartz, Gary T (Committee member) / Arizona State University (Publisher)
Created2012
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Description
Cercopithecid primates today occupy the greatest geographic and climatic range of any non-human primate group. Pliocene and Pleistocene cercopithecids are often found together in fossil deposits across East and South Africa, raising the question of how these species co-occurred with one another and survived in increasingly arid and seasonal environments.

Cercopithecid primates today occupy the greatest geographic and climatic range of any non-human primate group. Pliocene and Pleistocene cercopithecids are often found together in fossil deposits across East and South Africa, raising the question of how these species co-occurred with one another and survived in increasingly arid and seasonal environments. Aspects of shearing ability, molar enamel thickness, and relative incisor, premolar, and molar proportions were analyzed in principal component analysis and used to generate six potential models of the cercopithecid dental morphological niche. Resulting principal component axes distinguish between taxa with varying proportions of leaves, fruit, insects, and seeds in the diet, but lose some clarity when variable subsets are used that exclude poorly-preserved or wear-restricted variables. Resampling was used to reconstruct the aggregate dental morphological niches of cercopithecid communities (taxocenes) from Africa and Asia today and from the African Pliocene and Pleistocene. Modern Asian cercopithecid taxocenes occupy a more restricted niche than their counterparts in Africa, but in both regions variation in taxocene structure is linked with past and current climate factors related to precipitation, temperature, and seasonality. Fossil cercopithecids from the Turkana Basin occupy an expanded niche in comparison to modern African and Asian taxocenes. In contrast, South African fossil taxocenes occupy a more distinct and restricted niche, which may reflect a mix of paleoenvironmental and taphonomic factors. Overall these results are consistent with existing research on modern African and Asian primate taxocene diversity and highlight the utility of a dental metric model for examining community evolution among Plio-Pleistocene cercopithecids in Africa. Evidence for a possible niche expansion during the early Pleistocene coincides with a period of well-documented hominin co-occurrence at the same fossil sites, suggesting that these two primate groups were diversifying in response to shared environmental stimuli.
ContributorsSmail, Irene (Author) / Reed, Kaye E (Thesis advisor) / Campisano, Christopher J (Committee member) / Gilbert, Christopher C (Committee member) / Arizona State University (Publisher)
Created2021
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Description
Age is a key axis upon which social identities and social relationships are negotiated over the life course, and early life experiences can also have significant implications for individual and population health outcomes. However, children and childhood have historically been marginalized in the study of past societies, and non-elite children

Age is a key axis upon which social identities and social relationships are negotiated over the life course, and early life experiences can also have significant implications for individual and population health outcomes. However, children and childhood have historically been marginalized in the study of past societies, and non-elite children have been remarkably invisible in reports on ancient Greece. This dissertation employs a bioarchaeological approach to investigate age-related social identities, early childhood health, and the impact of prolonged childhood illness on familial social dynamics during the Archaic Period in Athens, Greece (ca. 700-480 BCE), focusing on 179 preadults interred at the non-elite cemetery of Phaleron. First, contextual mortuary evidence is used to investigate how age-at-death influenced burial at Phaleron, revealing insights into the timing of the personhood acquisition, age-related social transitions, and individual agency in burial practice as expressed through variation in mortuary treatment. Then paleopathological analysis of preadult skeletal remains is leveraged to investigate early childhood health outcomes, demonstrating that children at Phaleron experienced early life physiological stress, including nutritional insufficiency that may be linked to maternal health. Furthermore, evidence of poor health among non-survivors is argued to have significant implications for later life health among those who survived to adulthood. Finally, sociohistorical, contextual, and paleopathological data are synthesized to investigate the social implications of healthcare at Phaleron. The results of this multi-scalar analysis indicate that children interred at Phaleron not only survived extended periods of potentially debilitating illness, but also that their survival would not have been possible without a community of caregivers. Moreover, the age at which children experienced illness would have significantly impacted the types of healthcare needed and the burdens that care would have placed on the household. This dissertation demonstrates the promise of early childhood health and social identity as subjects of bioarchaeological inquiry in ancient Greece and underscores the social and emotional impacts of childcare and loss on the communities that buried their deceased at Phaleron. Consequently, it lays the groundwork for future research on children and childhood in ancient Greece and the study of past lifeways in Archaic Athens.
ContributorsRothwell, Jessica E. (Author) / Buikstra, Jane E. (Thesis advisor) / Knudson, Kelly J. (Committee member) / Stojanowski, Christopher M. (Committee member) / Anderson, Greg (Committee member) / Arizona State University (Publisher)
Created2024
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This dissertation research describes the hunting behavior of early modern humans through the analysis of vertebrate faunal remains from Contrebandiers Cave, Morocco. Contrebandiers Cave is located in the town of Témara and is roughly 250 meters from the current shoreline of the Atlantic Ocean. The cave was excavated in the

This dissertation research describes the hunting behavior of early modern humans through the analysis of vertebrate faunal remains from Contrebandiers Cave, Morocco. Contrebandiers Cave is located in the town of Témara and is roughly 250 meters from the current shoreline of the Atlantic Ocean. The cave was excavated in the 1950s and 1970s by l’Abbé Roche, and again starting in 2007 by Dibble and El Hajraoui with total station plotting of finds. Contrebandiers Cave contains Middle Stone Age (MSA) deposits dated to Marine Isotope Stages (MIS) 5e, 5d and 5c, ~120,000 to ~96,000 years ago. The Later Stone Age (LSA) deposits are dated to MIS 2, ~20,000 years ago. The entirety of the ~12,000 vertebrate faunal remains from Dibble and El Hajraoui’s excavation were analyzed for taxonomic and taphonomic identification.

A total of 67 vertebrate taxa were identified and include ungulates, carnivores, lagomorphs, birds, tortoises, snakes and fish. The faunal remains from Contrebandiers Cave preserve surface modification that indicates both humans and carnivores acted as agents of prey accumulation. Skeletal element representation and surface modification of ungulate remains suggest that humans had primary access to small, medium and large-bodied prey. In the MSA levels, carnivore skeletal remains preserve surface modification that is interpreted as being indicative of behavior associated with skinning for fur removal.

The vertebrate faunal remains from MIS 5e and 5d indicate that humans were hunting grazers and mixed feeders from open habitats and suids from mixed habitats. The faunal remains from MIS 5c indicate that humans focused less on suids and more on mixed feeders from open habitats. The vertebrate faunal remains from MIS 2 reveal humans hunting grazers from dry, open habitats. This research provides a description of human hunting behavior in North Africa, and contributes to our understanding of early modern human behavior prior to dispersal out of Africa.
ContributorsHallett, Emily Yuko (Author) / Marean, Curtis W (Thesis advisor) / Reed, Kaye E (Committee member) / Dibble, Harold L. (Committee member) / Arizona State University (Publisher)
Created2018
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Description
Social identities are fundamental to the way individuals and groups define themselves. Archaeological approaches to social identities in the Andes emphasize the importance of group identities such as ethnicity and community identity, but studies of gender and age identities are still uncommon. In this dissertation, I build on these earlier

Social identities are fundamental to the way individuals and groups define themselves. Archaeological approaches to social identities in the Andes emphasize the importance of group identities such as ethnicity and community identity, but studies of gender and age identities are still uncommon. In this dissertation, I build on these earlier approaches to Andean social identities and consider community, gender, and age identities at the site of Chiribaya Alta using case studies.

The coastal Ilo Chiribaya polity is associated with the Andean Late Intermediate Period in the lower Osmore drainage of southern Peru. Previous analyses indicate that Chiribaya sites in this area formed a señorío, an Andean chiefdom with separate occupational groups of fishers and farmers. The most complex excavated Chiribaya site in this region is Chiribaya Alta. At this time, excavations have sampled nine of the cemeteries present at the site. Two of these cemeteries, four and seven, have the most elaborate burials at the site and are each associated with different occupational communities.

This dissertation examines community, gender, and age identities at Chiribaya Alta through the use of three case studies. The first case study argues that the iconographic designs on coca bags interred with the dead signified occupational community identities. Coca bags buried in cemetery four have designs relating to mountains and farming, whereas those from cemetery seven have symbols associated with water. These designs correspond to the occupational community groups associated with each of these cemeteries. The second case study uses grave good presence and absence to examine the nature of gender roles and identity at Chiribaya Alta. Multiple correspondence analysis indicates that normative gender roles are reflected in grave good assemblages, but that gender identity was flexible at the individual level. The final case study presents newly generated age-at-death estimations using transition analysis combined with mortuary analyses to explore the manner in which gender and age intersect for older individuals at Chiribaya Alta. This final paper argues that there is an elderly identity present amongst individuals at Chiribaya Alta and that gender and age intersect to impact the lives of older men and women differently.
ContributorsSchach, Emily Ann (Author) / Buikstra, Jane E. (Thesis advisor) / Knudson, Kelly J. (Committee member) / Geller, Pamela L. (Committee member) / Arizona State University (Publisher)
Created2019
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Description
Providing an environmental context to early hominins is as important as describing the hominin fossils themselves, because evolutionary processes are tightly linked to everchanging ecosystems that vary across space and through time. An optimal understanding of ecosystems changes is critical to formulate and test hypotheses regarding human evolution and adaptation.

Providing an environmental context to early hominins is as important as describing the hominin fossils themselves, because evolutionary processes are tightly linked to everchanging ecosystems that vary across space and through time. An optimal understanding of ecosystems changes is critical to formulate and test hypotheses regarding human evolution and adaptation. Fortunately, the fossil record has yielded abundant remains of mammals which can be used to explore the possible causal relationships between environmental change and mammal – including hominin –evolution. Although many studies have already been conducted on this topic, most of them are framed at large spatial and temporal scales. Instead, this dissertation focuses on the evolution and paleoecology of only one group of mammals (the Suidae) in a specific geographical area (lower Awash Valley in Ethiopia) and within a constrained time frame (3.8–2.6 Ma). Three dissertation papers address: 1) changes in suid taxonomic composition in relation to Late Pliocene faunal turnover ~2.8 Ma in the Lee-Adoyta basin, Ledi-Geraru; 2) comparisons of suid diets from Hadar (~3.45–2.95 Ma) with respect to those of Kanapoi (~4.1 Ma, West Turkana, Kenya); 3) the dietary ecology of the suids from Woranso-Mille (~3.8–3.2 Ma). Results of these papers show that 1) after ~2.8 Ma there is a replacement of suid species that is coupled with low relative abundance of suids. This is compatible with more open and/or arid environments at this time; 2) suid dietary breadth was broader in Hadar than in Kanapoi, but this is mostly driven by the dietary niche space occupied by Kolpochoerus in Hadar, a suid genus absent from Kanapoi; 3) suid diets vary both temporally and geographically within the lower Awash Valley. Kolpochoerus incorporates more C4 resources (e.g., grasses) in its diet after ~3.5 Ma and in general, suids after ~3.5 Ma in Woranso-Mille had C4-enriched diets in comparison with those from nearby Hadar and Dikika. Presumably, the changes in suid communities (relative abundance and taxonomic composition) and dietary shifts observed in suids were triggered by climatic and habitat changes that also contributed to shape the behavioural and morphological evolution of early hominins.
ContributorsAguilar Lazagabaster, Ignacio (Author) / Reed, Kaye E (Thesis advisor) / Kimbel, William H. (Committee member) / Ungar, Peter S. (Committee member) / Arizona State University (Publisher)
Created2018
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This dissertation focuses on the diversity inherent to the process of social community construction. Building upon previous archaeological and bioarchaeological studies of community identities, the current project emphasizes the need for consideration of the impact of diversity on community identity formation in the past and illustrates the utility of a

This dissertation focuses on the diversity inherent to the process of social community construction. Building upon previous archaeological and bioarchaeological studies of community identities, the current project emphasizes the need for consideration of the impact of diversity on community identity formation in the past and illustrates the utility of a bioarchaeological approach for undertaking this task. Three specific aspects of community formation are addressed: (1) the relationship between symbolic community boundaries and geographic space, (2) the influence of diverse discourses of intra-community sub-groups on community formation, and (3) the negotiation of community boundaries by outsiders. To investigate these aspects of community construction in the past, dietary practices and mortuary rituals of the Late Intermediate Period (c. AD 900-1470) Ychsma society of the central Peruvian coast are examined as a case study. Previous anthropological and sociological studies demonstrate that diet and burial customs are common mechanisms used in processes of group identification around the world, including the Andes. In the current study, analyses of materials from Armatambo and Rinconada Alta in the Rimac Valley are used to examine the ways in which isotopic and dental indicators of diet and archaeological contextual indicators of mortuary rituals correspond with or crosscut spatial burial patterns and additional groups based on sex, age at death, and biogeochemically reconstructed residential origins. Observed patterns are interpreted using a theoretical framework that incorporates sociocultural theory of identity with pre-Columbian Andean ideology of the body, self, and social environment. Results reveal differences in large-scale trends in diet and mortuary practices associated with burial at each site that are interpreted as evidence of symbolic community boundaries between sites. Complexities within larger trends reveal evidence of internal diversity as well as fluidity across community boundaries. Specifically, evidence is presented for intra-community dietary differences, intra-community differences associated with age and sex, and finally evidence of external relationships. This consideration of diversity in community identity construction is concluded to profoundly refine current understandings of Ychsma social interactions. Consequently, this study demonstrates empirical investigation of social diversity is necessary for understanding the complex nature of the social construction of communities in the past.
ContributorsMarsteller, Sara Jane (Author) / Knudson, Kelly J. (Thesis advisor) / Buikstra, Jane E. (Committee member) / Scott, Rachel E. (Committee member) / Arizona State University (Publisher)
Created2015
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This dissertation addresses the role of kinship and residential mobility during the transition from Final Neolithic to Early Bronze Age (ca. 3500 – 2500 BC) in Attica, Greece. It examines descent systems, ancestor formation, and the interplay between biological, social, and spatial structure in mortuary practices. It also evaluates the

This dissertation addresses the role of kinship and residential mobility during the transition from Final Neolithic to Early Bronze Age (ca. 3500 – 2500 BC) in Attica, Greece. It examines descent systems, ancestor formation, and the interplay between biological, social, and spatial structure in mortuary practices. It also evaluates the nature and degree of residential mobility and its potential role in the formation and maintenance of social networks. Archaeological hypotheses on the kin-based structure of formal cemeteries, the familial use of collective tombs, marriage practices and mate exchange, and relocation were tested focusing on the Early Helladic cemetery of Tsepi at Marathon. Tsepi constitutes the earliest formally organized cemetery on the Greek mainland and it has also contributed to enduring debates over the nature of the interaction between the eastern Attic coast and the central Aegean islands.

This study integrates osteological, biogeochemical, and archaeological data. Inherited dental and cranial features were used to examine biological relatedness and postmarital residence (biodistance analysis). Biochemical analysis of archaeological and modern samples was conducted to examine the geographic origins of the individuals buried in the cemetery and reconstruct mobility patterns. Osteological and biogeochemical data were interpreted in conjunction with archaeological and ethnographic/ethnohistoric data.

The results generally supported a relationship between spatial organization and biological relatedness based on phenotypic similarity at Tsepi. Postmarital residence analysis showed exogamous practices and tentatively supported higher male than female mobility. This practice, along with dietary inferences, could also be suggestive of maritime activities. Biogeochemical analysis showed a local character for the cemetery sample (96%). The common provenance of the three non-local individuals might reflect a link between Tsepi and a single locale. Burial location was not determined by provenance or solely by biological relatedness. Overall, the results point towards more nuanced reconstructions of mobility in prehistoric Aegean and suggest that burial location depended on a complex set of inter-individual relationships and collective identities. The contextualized bioarchaeological approach applied in this study added to the anthropological investigations of social practices such as kin relations (e.g., biological, marital, social kinship) and residential relocation as diachronic mechanisms of integration, adaptation, or differentiation.
ContributorsPrevedorou, Eleni Anna (Author) / Buikstra, Jane E. (Thesis advisor) / Knudson, Kelly J. (Committee member) / Stojanowski, Chris M (Committee member) / Fox, Sherry C. (Committee member) / Rutter, Jeremy B. (Committee member) / Arizona State University (Publisher)
Created2015
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This study explores how early modern humans used stone tool technology to adapt to changing climates and coastlines in the Middle Stone Age of South Africa. The MSA is associated with the earliest fossil evidence for modern humans and complex cultural behaviors during a time period of dramatic climate change.

This study explores how early modern humans used stone tool technology to adapt to changing climates and coastlines in the Middle Stone Age of South Africa. The MSA is associated with the earliest fossil evidence for modern humans and complex cultural behaviors during a time period of dramatic climate change. Human culture allows for the creation, use, and transmission of technological knowledge that can evolve with changing environmental conditions. Understanding the interactions between technology and the environment is essential to illuminating the role of culture during the origin of our species. This study is focused on understanding ancient tool use from the study of lithic edge damage patterns at archaeological assemblages in southern Africa by using image-based quantitative methods for analyzing stone tools. An extensive experimental program using replicated stone tools provides the comparative linkages between the archaeological artifacts and the tasks for which they were used. MSA foragers structured their tool use and discard behaviors on the landscape in several ways – by using and discarding hunting tools more frequently in the field rather than in caves/rockshelters, but similarly in coastal and interior contexts. This study provides evidence that during a significant microlithic technological shift seen in southern Africa at ~75,000 years ago, new technologies were developed alongside rather than replacing existing technologies. These results are compared with aspects of the European archaeological record at this time to identify features of early human technological behavior that may be unique to the evolutionary history of our species.
ContributorsSchoville, Benjamin J (Author) / Marean, Curtis W (Thesis advisor) / Barton, Michael (Committee member) / Hill, Kim (Committee member) / Arizona State University (Publisher)
Created2016