Glen Canyon Dam Adaptive Management Program Administrative History

While the ecology and evolution of partial migratory systems (defined broadly to include skip spawning) have been well studied, we are only beginning to under- stand how partial migratory populations are responding to ongoing environmen- tal change. Environmental change can lead to differences in the fitness of residents and migrants, which could eventually lead to changes in the frequency of the strategies in the overall population. Here, we address questions concerning the life history of the endangered Gila cypha (humpback chub) in the regulated Colorado River and the unregulated tributary and primary spawning area, the Little Colorado River. We develop eight multistate models for the population based on three movement hypotheses, in which states are defined in terms of fish size classes and river locations. We fit these models to mark–recapture data col- lected in 2009–2012. We compare survival and growth estimates between the Col- orado River and Little Colorado River and calculate abundances for all size classes. The best model supports the hypotheses that larger adults spawn more frequently than smaller adults, that there are residents in the spawning grounds, and that juveniles move out of the Little Colorado River in large numbers during the monsoon season (July–September). Monthly survival rates for G. cypha in the Colorado River are higher than in the Little Colorado River in all size classes; however, growth is slower. While the hypothetical life histories of life-long resi- dents in the Little Colorado River and partial migrants spending most of its time in the Colorado River are very different, they lead to roughly similar fitness expectations when we used expected number of spawns as a proxy. However, more research is needed because our study period covers a period of years when conditions in the Colorado River for G. cypha are likely to have been better than has been typical over the last few decades.

Between 1999 and 2005, drought in the western United States led to a >44 m fall in the level of Lake Powell (Arizona-Utah), the nation's second-largest reservoir. River discharges to the reservoir were halved, yet the rivers still incised the tops of deltas left exposed along the rim of the reservoir by the lake-level fall. Erosion of the deltas enriched the rivers in sediment such that upon entering the reservoir they discharged plunging subaqueous gravity flows, one of which was imaged acoustically. Repeat bathymetric surveys of the reservoir show that the gravity flows overtopped rockfalls and formed small subaqueous fans, locally raising sediment accumulation rates 10–100-fold. The timing of deep-basin deposition differed regionally across the reservoir with respect to lake-level change. Total mass of sediment transferred from the lake perimeter to its bottom equates to ~22 yr of river input.