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Description
The ability of Neandertals to cope with the oscillating climate of the late Pleistocene and the extent to which these climate changes affected local Neandertal habitats remain unanswered anthropological topics of considerable scientific interest. Understanding the impact of climatic instability on Neandertals is critical for reconstructing the behaviors of our

The ability of Neandertals to cope with the oscillating climate of the late Pleistocene and the extent to which these climate changes affected local Neandertal habitats remain unanswered anthropological topics of considerable scientific interest. Understanding the impact of climatic instability on Neandertals is critical for reconstructing the behaviors of our closest fossil relatives and possibly identifying factors that contributed to their extinction. My work aimed to test the hypotheses that 1) cold climates stressed Neandertal populations, and 2) that global climate changes affected local Neandertal habitats. An analysis of Neandertal butchering on Cervus elaphus, Rangifer tarandus, and Capreolus capreolus skeletal material deposited during global warm and cold phases from two French sites - Pech de l'Azé IV and Roc de Marsal - was conducted to assess the impact of climate change on butchering strategies and resource extraction. Results from a statistical analysis of surface modification on all marrow yielding long bones, including the 1st phalanx, demonstrated that specimens excavated from the cold levels at each cave have more cut marks (Wald χ2= 51.33, p= <0.001) and percussion marks (Wald χ2= 4.92, p= 0.02) than specimens from the warm levels after controlling for fragment size. These results support the hypothesis that Neandertals were nutritionally stressed during glacial cycles. The hypothesis that global climates affected local habitats was tested through radiogenic strontium isotopic reconstruction of large herbivore mobility patterns (e.g., Bison, Equus, Cervus and Rangifer), because it is known that in the northern hemisphere, mammals migrate less in warm, well-vegetated environments, but more in cold, open environments. Identifying isotopic variation in mammalian fossils enables mobility patterns to be inferred, providing an indication of whether environments at Pech de l'Azé IV and Roc de Marsal tracked global climates. Results from this study indicate that Neandertal prey species within the Dordogne Valley of France did not undertake long distance round-trip migrations in glacial or interglacial cycles, maintaining the possibility that local habitats did not change in differing climatic cycles. However, because Neandertals were nutritionally stressed the most likely conclusion is that glacial cycles decreased herbivore populations, thus stressing Neandertals.
ContributorsHodgkins, Jamie Melichar (Author) / Marean, Curtis W (Thesis advisor) / Reed, Kaye E (Thesis advisor) / Knudson, Kelly J. (Committee member) / Spencer, Lillian M (Committee member) / Arizona State University (Publisher)
Created2012
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Description
Sedimentary basins in the Afar Depression, Ethiopia archive the progression of continental breakup, record regional changes in east African climate and volcanism, and host what are arguably the most important fossiliferous strata for studying early human evolution and innovation. Significant changes in rift tectonics, climate, and faunal assemblages occur between

Sedimentary basins in the Afar Depression, Ethiopia archive the progression of continental breakup, record regional changes in east African climate and volcanism, and host what are arguably the most important fossiliferous strata for studying early human evolution and innovation. Significant changes in rift tectonics, climate, and faunal assemblages occur between 3-2.5 million years ago (Ma), but sediments spanning this time period are sparse. In this dissertation, I present the results of a geologic investigation targeting sediments between 3-2.5 Ma in the central and eastern Ledi Geraru (CLG and ELG) field areas in the lower Awash Valley, using a combination of geologic mapping, stratigraphy, and tephra chemistry and dating. At Gulfaytu in CLG, I mapped the northern-most outcrops of the hominin-bearing Hadar Formation (3.8-2.9 Ma), a 20 m-thick section of flat-lying lacustrine sediments containing 8 new tephras that directly overlie the widespread BKT-2 marker beds (2.95 Ma). Paleolake Hadar persisted after 2.95 Ma, and the presence and characteristics of the Busidima Formation (2.7-0.016 Ma) indicates Gulfaytu was affected by a reversal in depositional basin polarity. Combined with regional and geophysical data, I show the Hadar Formation underlying CLG is >300 m thick, supporting the hypothesis that it was the lower Awash Pliocene depocenter. At ELG, I mapped >300 m of sediments spanning 3.0-2.45 Ma. These sediments coarsen upward and show a progression from fluctuating lake conditions to fluvial landscapes and widespread soil development. This is consistent with the temporal change in depositional environments observed elsewhere in the lower Awash Valley, and suggests that these strata are correlative with the Hadar Formation. Furthermore, the strata and basalts at ELG are highly faulted, and overprinted by shifting extension directions attributed to the northern migration of the Afar triple junction. The presence of fossiliferous beds and stone tools makes ELG a high-priority target for anthropological and archaeological research. This study provides a new temporally-calibrated and high-resolution record of deposition, volcanism, and faulting patterns during a period of significant change in the Afar.
ContributorsDiMaggio, Erin Nicole (Author) / Arrowsmith, J Ramon (Thesis advisor) / Whipple, Kelin X (Committee member) / Heimsath, Arjun M (Committee member) / Clarke, Amanda B (Committee member) / Reed, Kaye E (Committee member) / Arizona State University (Publisher)
Created2013
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Description
Identifying the ecological role, or niche, that a species occupies within their larger community elucidates environmental adaptability and evolutionary success. This dissertation investigates the occupied niche of chimpanzees (Pan troglodytes schweinfurthii) living in an open, dry savanna-woodland environment by examining patterns of resource use and interspecific interactions. Data were collected

Identifying the ecological role, or niche, that a species occupies within their larger community elucidates environmental adaptability and evolutionary success. This dissertation investigates the occupied niche of chimpanzees (Pan troglodytes schweinfurthii) living in an open, dry savanna-woodland environment by examining patterns of resource use and interspecific interactions. Data were collected October 2010--November 2011 at Issa, in the Ugalla region of western Tanzania, which is one of the driest, most open, and seasonal habitats inhabited by chimpanzees. Unlike most primatological studies which employ methods that include focal follows, this study focused instead on observing 'resource patches' for chimpanzees. Patch focals allow for the observation of all animals within a study area; capture resources that are not used by the study species; and are particularly well suited for unhabituated communities. In order to better understand relationships between environment and behavior, data collected at Issa are compared with published data from other chimpanzee populations. Issa chimpanzees were expected to have broader resource use than forest chimpanzees, as well as increased competition with other fauna, due to fewer available resources. However, in contrast to the assumption of food scarcity in dry habitats, dietary resources were available throughout the year. Like other populations, the diet of Issa chimpanzees consisted of mostly fruit, but unlike at other sites, the majority of plants consumed were woodland species. Additionally, although chimpanzees and other fauna shared spatial and dietary resources, there was only nominal overlap. These results point to extremely low levels of indirect competition between chimpanzees and other fauna. Despite extensive study of forest chimpanzees, little is known about their role within their faunal community in open, dry habitats, nor about how greater seasonality affects resource use. This project addresses both of these important issues and fosters novel approaches in anthropological studies, especially in reference to chimpanzee ecology and evolution. Understanding current chimpanzee behavioral relationships with their environments shapes hypotheses about their pasts, and also informs predictions about behaviors of similar taxa in paleo-environments. Lastly, examining the ecological role of chimpanzees within their larger communities will influence the formation of, as well as evaluate, conservation strategies.
ContributorsRussak, Samantha M (Author) / Reed, Kaye E (Thesis advisor) / Nash, Leanne T. (Committee member) / Schwartz, Gary T (Committee member) / Arizona State University (Publisher)
Created2013
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Description
The earliest Eocene marked the appearance of the first North American euprimates (adapids, omomyids). Despite the fact that leading hypotheses assert that traits involved in food acquisition underlie euprimate origination and early diversification, the precise role that dietary competition played in establishing euprimates as successful members of mammalian communities is

The earliest Eocene marked the appearance of the first North American euprimates (adapids, omomyids). Despite the fact that leading hypotheses assert that traits involved in food acquisition underlie euprimate origination and early diversification, the precise role that dietary competition played in establishing euprimates as successful members of mammalian communities is unclear. This is because the degree of niche overlap between euprimates and all likely mammalian dietary competitors ("the euprimate competitive guild") is unknown. This research determined which of three major competition hypotheses - non-competition, strong competition, and weak competition - characterized the late Paleocene-early Eocene euprimate competitive guild. Each of these hypotheses is defined by a unique temporal pattern of niche overlap between euprimates and their non-euprimate competitors, allowing an evaluation of the nature of dietary competitive interactions surrounding the earliest euprimates in North America. Dietary niches were reconstructed for taxa within the fossil euprimate competitive guild using molar morphological measures determined to discriminate dietary regimes in two extant mammalian guilds. The degree of dietary niche separation among taxa was then evaluated across a series of fossil samples from the Bighorn Basin, Wyoming just prior to, during, and after euprimate origination. Statistical overlap between each pair of euprimate and non-euprimate dietary niches was determined using modified multivariate pairwise comparisons using distances in a multidimensional principal component "niche" space. Results indicate that euprimate origination and diversification in North America was generally characterized by the absence of dietary competition. This lack of competition with non-euprimates is consistent with an increase in the abundance and diversity of euprimates during the early Eocene, signifying that the "success" of euprimates may not be the result of direct biotic interactions between euprimates and other mammals. An examination of the euprimate dietary niche itself determined that adapids and omomyids occupied distinct niches and did not engage in dietary competition during the early Eocene. Furthermore, changes in euprimate dietary niche size over time parallel major climatic shifts. Reconstructing how both biotic and abiotic mechanisms affected Eocene euprimates has the potential to enhance our understanding of these influences on modern primate communities.
ContributorsStroik, Laura (Author) / Schwart, Gary T (Thesis advisor) / Reed, Kaye E (Committee member) / Campisano, Christopher J (Committee member) / Gunnell, Gregg F. (Committee member) / Arizona State University (Publisher)
Created2014