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This dissertation research investigates both spatial and temporal aspects of Bronze Age land use and land cover in the Eastern Mediterranean using botanical macrofossils of charcoal and charred seeds as sources of proxy data. Comparisons through time and over space using seed and charcoal densities, seed to charcoal ratios, and

This dissertation research investigates both spatial and temporal aspects of Bronze Age land use and land cover in the Eastern Mediterranean using botanical macrofossils of charcoal and charred seeds as sources of proxy data. Comparisons through time and over space using seed and charcoal densities, seed to charcoal ratios, and seed and charcoal identifications provide a comprehensive view of island vs. mainland vegetative trajectories through the critical 1000 year time period from 2500 BC to 1500 BC of both climatic fluctuation and significant anthropogenic forces. This research focuses particularly on the Mediterranean island of Cyprus during this crucial interface of climatic and human impacts on the landscape. Macrobotanical data often are interpreted locally in reference to a specific site, whereas this research draws spatial comparisons between contemporaneous archaeological sites as well as temporal comparisons between non-contemporaneous sites. This larger perspective is particularly crucial on Cyprus, where field scientists commonly assume that botanical macrofossils are poorly preserved, thus unnecessarily limiting their use as an interpretive proxy. These data reveal very minor anthropogenic landscape changes on the island of Cyprus compared to those associated with contemporaneous mainland sites. These data also reveal that climatic forces influenced land use decisions on the mainland sites, and provides crucial evidence pertaining to the rise of early anthropogenic landscapes and urbanized civilization.
ContributorsKlinge, JoAnna M (Author) / Fall, Patricia L. (Thesis advisor) / Falconer, Steven E. (Committee member) / Brazel, Anthony J. (Committee member) / Pigg, Kathleen B (Committee member) / Arizona State University (Publisher)
Created2013
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Description
Acceptance of the plant group Martyniaceae as a distinct family has long been questioned. Previously placed in the family Pedaliaceae, the Martyniaceae have been allied to numerous other families within the order Lamiales. The objectives of this study include the investigation of the placement of the Martyniaceae within the order

Acceptance of the plant group Martyniaceae as a distinct family has long been questioned. Previously placed in the family Pedaliaceae, the Martyniaceae have been allied to numerous other families within the order Lamiales. The objectives of this study include the investigation of the placement of the Martyniaceae within the order Lamiales using molecular data (chloroplast DNA sequences), the further examination of the internal relationships of the Martyniaceae using an expanded nuclear and chloroplast sequences data set, and the construction of a taxonomic treatment of the family that includes all published names and taxa in the Martyniaceae. An analysis of the Lamiales using two chloroplast gene regions (ndhF and rps16) reveals that the Martyniaceae should be segregated from the family Pedaliaceae, but is not able to support the placement of any of its putatively-related families as sister to the Martyniaceae. Sequences from 151 taxa of the Lamiales are included in the analysis, including six representatives from the Martyniaceae. An analysis of the Martyniaceae using three chloroplast gene regions (psbA-trnH spacer, trnQ-5'rps16 intergenic spacer, and trnS-trnG-trnG spacer and intron) and the Internal Transcribed Spacer resolves two major clades within the Martyniaceae corresponding to the North American taxa (Martynia and Proboscidea) and the South American taxa (Craniolaria, Holoregmia, and Ibicella). Sequences from all five genera and 15 taxa were included in the analysis. Results from the molecular phylogenetic analyses are incorporated into a revised taxonomic treatment of the family. Five genera and thirteen species are recognized for the family Martyniaceae.
ContributorsGutiérrez, Raúl (Author) / Wojciechowski, Martin F (Thesis advisor) / Pigg, Kathleen B (Committee member) / Landrum, Leslie R (Committee member) / Butterworth, Charlie (Committee member) / Arizona State University (Publisher)
Created2011
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Description
The Juglandaceae (walnuts, hickories, pecans) has one of the best-documented fossil records in the Northern Hemisphere. The oldest modern genus, Cyclocarya, today restricted to China, first appears in the late Paleocene (57 ma) of North Dakota, USA. Unlike walnuts and pecans that produce edible fruits dispersed by mammals, Cyclocarya fruits

The Juglandaceae (walnuts, hickories, pecans) has one of the best-documented fossil records in the Northern Hemisphere. The oldest modern genus, Cyclocarya, today restricted to China, first appears in the late Paleocene (57 ma) of North Dakota, USA. Unlike walnuts and pecans that produce edible fruits dispersed by mammals, Cyclocarya fruits are small nutlets surrounded by a prominent circular wing, and are thought to be wind- or water-dispersed. The current study provides the first evidence that fossil fruits were different from modern forms in the number and organization of their attachment to reproductive branches, and in their anatomical structure. Unlike the modern genus that bears separate pistillate and staminate flowers the fossil fruits had attached pollen-bearing structures. Unisexual pollen catkins are also present, suggesting the fossil Cyclocarya may have differed from its modern relative in this feature. Like several other plants from the late Paleocene Almont/Beicegel Creek floras, Cyclocarya shows a mosaic combination of characters not seen in their modern counterparts. Fossils were collected from the field, and examined for specimens exposed on the weathered rock surface. Specimens from Almont were photographed with reflected light, while those from Beicegel Creek cut into slabs and prepared by etching the rock matrix in 49% hydrofluoric and re-embedding the exposed plant material in cellulose acetate and acetone to make "peels". Selected specimens are cut out, mounted on microscope slides, and studied with light microscopy. These fossil fruits were studied because they are the earliest fossil evidence of Cyclocarya. They are exceptionally preserved and thus provide critical structural evidence for changes in that occurred during the evolution of plants within this lineage. Because Cyclocarya fruits are winged, they might be assumed to be wind-dispersed. Their radial symmetry does not have the aerodynamic qualities typical of wind-dispersed fruits, and may have been dispersed by water.
ContributorsTaylor, Malcom DeWitt (Author) / Pigg, Kathleen B (Thesis advisor) / Wojciechowski, Martin F (Committee member) / Devore, Melanie L (Committee member) / Farmer, Jack (Committee member) / Gill, Anthony (Committee member) / Arizona State University (Publisher)
Created2010