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Description
This dissertation shows that the central conceptual feature and explanatory motivation of theories of evolutionary directionality between 1890 and 1926 was as follows: morphological variation in the developing organism limits the possible outcomes of evolution in definite directions. Put broadly, these theories maintained a conceptual connection between development and evolution as inextricably associated phenomena. This project develops three case studies. The first addresses the Swiss-German zoologist Theodor Eimer's book Organic Evolution (1890), which sought to undermine the work of noted evolutionist August Weismann. Second, the American paleontologist Edward Drinker Cope's Primary Factors (1896) developed a sophisticated system of inheritance that included the material of heredity and the energy needed to induce and modify ontogenetic phenomena. Third, the Russian biogeographer Leo Berg's Nomogenesis (1926) argued that the biological world is deeply structured in a way that prevents changes to morphology taking place in more than one or a few directions. These authors based their ideas on extensive empirical evidence of long-term evolutionary trajectories. They also sought to synthesize knowledge from a wide range of studies and proposed causes of evolution and development within a unified causal framework based on laws of evolution. While being mindful of the variation between these three theories, this project advances "Definitely Directed Evolution" as a term to designate these shared features. The conceptual coherence and reception of these theories shows that Definitely Directed Evolution from 1890 to 1926 is an important piece in reconstructing the wider history of theories of evolutionary directionality.
ContributorsUlett, Mark Andrew (Author) / Laubichler, Manfred D (Thesis advisor) / Hall, Brian K (Committee member) / Lynch, John (Committee member) / Maienschein, Jane (Committee member) / Smocovitis, Vassiliki B (Committee member) / Arizona State University (Publisher)
Created2014
Description
During the 1960s, the long-standing idea that traits or behaviors could be
explained by natural selection acting on traits that persisted "for the good of the group" prompted a series of debates about group-level selection and the effectiveness with which natural selection could act at or across multiple levels of biological organization. For some this topic remains contentious, while others consider the debate settled, even while disagreeing about when and how resolution occurred, raising the question: "Why have these debates continued?"
Here I explore the biology, history, and philosophy of the possibility of natural selection operating at levels of biological organization other than the organism by focusing on debates about group-level selection that have occurred since the 1960s. In particular, I use experimental, historical, and synthetic methods to review how the debates have changed, and whether different uses of the same words and concepts can lead to different interpretations of the same experimental data.
I begin with the results of a group-selection experiment I conducted using the parasitoid wasp Nasonia, and discuss how the interpretation depends on how one conceives of and defines a "group." Then I review the history of the group selection controversy and argue that this history is best interpreted as multiple, interrelated debates rather than a single continuous debate. Furthermore, I show how the aspects of these debates that have changed the most are related to theoretical content and empirical data, while disputes related to methods remain largely unchanged. Synthesizing this material, I distinguish four different "approaches" to the study of multilevel selection based on the questions and methods used by researchers, and I use the results of the Nasonia experiment to discuss how each approach can lead to different interpretations of the same experimental data. I argue that this realization can help to explain why debates about group and multilevel selection have persisted for nearly sixty years. Finally, the conclusions of this dissertation apply beyond evolutionary biology by providing an illustration of how key concepts can change over time, and how failing to appreciate this fact can lead to ongoing controversy within a scientific field.
explained by natural selection acting on traits that persisted "for the good of the group" prompted a series of debates about group-level selection and the effectiveness with which natural selection could act at or across multiple levels of biological organization. For some this topic remains contentious, while others consider the debate settled, even while disagreeing about when and how resolution occurred, raising the question: "Why have these debates continued?"
Here I explore the biology, history, and philosophy of the possibility of natural selection operating at levels of biological organization other than the organism by focusing on debates about group-level selection that have occurred since the 1960s. In particular, I use experimental, historical, and synthetic methods to review how the debates have changed, and whether different uses of the same words and concepts can lead to different interpretations of the same experimental data.
I begin with the results of a group-selection experiment I conducted using the parasitoid wasp Nasonia, and discuss how the interpretation depends on how one conceives of and defines a "group." Then I review the history of the group selection controversy and argue that this history is best interpreted as multiple, interrelated debates rather than a single continuous debate. Furthermore, I show how the aspects of these debates that have changed the most are related to theoretical content and empirical data, while disputes related to methods remain largely unchanged. Synthesizing this material, I distinguish four different "approaches" to the study of multilevel selection based on the questions and methods used by researchers, and I use the results of the Nasonia experiment to discuss how each approach can lead to different interpretations of the same experimental data. I argue that this realization can help to explain why debates about group and multilevel selection have persisted for nearly sixty years. Finally, the conclusions of this dissertation apply beyond evolutionary biology by providing an illustration of how key concepts can change over time, and how failing to appreciate this fact can lead to ongoing controversy within a scientific field.
ContributorsDimond, Christopher C (Author) / Collins, James P. (Thesis advisor) / Gadau, Juergen (Committee member) / Laubichler, Manfred (Committee member) / Armendt, Brad (Committee member) / Lynch, John (Committee member) / Arizona State University (Publisher)
Created2014
Description
Within the vast array of classical literature, many authors still have not been thoroughly examined. Among these, Palladas of Alexandria, a prominent epigrammatist in the Palatine Anthology, seems to import Epicurean themes. Based on the content and motifs of his poetry, I argue that Palladas leaned on Epicurean philosophy as presented in Lucretius's De Rerum Natura to salvage pagan identity in the midst of Christianity's increased popularity.
ContributorsNikpour, Rodmanned Arya (Author) / Tueller, Michael (Thesis director) / Lynch, John (Committee member) / Bing, Peter (Committee member) / Barrett, The Honors College (Contributor) / School of International Letters and Cultures (Contributor) / School of Historical, Philosophical and Religious Studies (Contributor)
Created2014-05
Description
In this thesis, I will be discussing the similarities between Stoicism (as both an ancient Greek and Roman philosophy, as well as how it is interpreted in the modern age) and modern therapeutic methods; However, I will not be developing any type of novel theory as to how Stoicism can be used as one of those therapeutic methods by itself. That would require a degree of psychological and medical knowledge that I, as an undergraduate student, do not yet possess and do not have the authority to expand upon in a safe manner. What the goal of this thesis is, instead, is to draw and explore parallels between the ideals and principles of stoicism (such as eudaimonia, ethics, and relative asceticism) as compared to modern therapeutic techniques, like cognitive-behavioral and dialectical-behavioral therapies. I will draw direct parallels between Stoic philosophy and the therapeutic treatments commonly used to address the symptoms of two psychiatric issues (Bipolar Mood Disorder and Obsessive Compulsive Disorder). I will also be addressing a third psychiatric case study, as relating to Stoic philosophy - suicide, and how our view of it has changed and progressed,both through a Stoic lens as well as from a contemporary psychological viewpoint.
As a result of drawing these parallels, this thesis will also explore some of the more modern uses of Stoicism - for example, those discussed in A Guide To The Good Life by William B Irvine, and Stoic Warriors by Nancy Sherman. Irvine focuses primarily on the use of Stoicism to avoid the factors of“chronic dissatisfaction” that afflict much of our modern-day lives - an absence of control, unhappiness, and erroneous personal values, to name a few. Sherman takes a more targeted approach - the application of Stoic philosophy to the workings of the military mentality and instinct. Sherman explores how being “Stoic” is taught as a part of military bearing, specifically when serving in the American forces. Stoic values are used to create a culture of discipline and self-control in the military - as Sherman puts it, “The idea that one’s happiness could depend solely on one’s own virtue, and that one’s agency and control might be bulletproof, appealed to [them]” (Sherman, 11). These authors’ perspectives are just two examples of how Stoicism can be applied in the modern age, as will be shown in further detail in subsequent sections.
As a result of drawing these parallels, this thesis will also explore some of the more modern uses of Stoicism - for example, those discussed in A Guide To The Good Life by William B Irvine, and Stoic Warriors by Nancy Sherman. Irvine focuses primarily on the use of Stoicism to avoid the factors of“chronic dissatisfaction” that afflict much of our modern-day lives - an absence of control, unhappiness, and erroneous personal values, to name a few. Sherman takes a more targeted approach - the application of Stoic philosophy to the workings of the military mentality and instinct. Sherman explores how being “Stoic” is taught as a part of military bearing, specifically when serving in the American forces. Stoic values are used to create a culture of discipline and self-control in the military - as Sherman puts it, “The idea that one’s happiness could depend solely on one’s own virtue, and that one’s agency and control might be bulletproof, appealed to [them]” (Sherman, 11). These authors’ perspectives are just two examples of how Stoicism can be applied in the modern age, as will be shown in further detail in subsequent sections.
ContributorsSimpson, Naomi (Author) / Lynch, John (Thesis director) / O'Neill, Joseph (Committee member) / School of Music (Contributor) / School of Social and Behavioral Sciences (Contributor) / Barrett, The Honors College (Contributor)
Created2020-05